Query Sequence
Gene or Gene Model ID
(e.g.: DDB_G0275689, DDB0214814)
BLAST Program
BLAST Database

DUST filter for BLASTN, SEG filter for all others.
Because of resource limits, when BLASTing against the Chromosomal database with filtering off, the filtering is actually set to 'm D' which turns off filtering for extending hits, but leaves filtering on when building the initial words.